Managed vs. unmanaged. Structure of beech forest stands (Fagus sylvatica L.) after 50 years of development, Central Bohemia
Abstract
In this study, the structure and spatial patterns of adjacent beech (Fagus sylvatica L.) forest stands, both those having undergone intensive management and those without management intervention over the last 50 years were analyzed. Data was collected from six 1ha permanent research plots. Four plots were established in varying parts of the reserve, comprising entirely of even-aged managed forest stands; two 1-ha permanent plots were located in a non-interventional stand. For each woody stem, basic dendrometrical data and coordinates were measured. The total volume of dead wood (standing and fallen) and its classes of decay were recorded. The structure and species diversity of forest stands were analyzed using skewness and the coefficient of variation of dbh distribution, the Gini index, the Shannon index, the mingling index, the Pielou index of segregation and the Hopkins-Skellam, Pielou-Mountford, Clark-Evans aggregation indices, the diameter differentiation index and Ripley K function. The diameter distribution of the managed stands was generally normal, albeit with a large diameter range and with a typical flat diameter curve. A general lack of giant trees, lower rates of coarse woody debris (from 0.2 to 0.5% of the total living volume) and random tree distribution were also confirmed. Within the investigated area, the diameter differentiation and Gini indices along with the degree of skewness and the coefficient of variation made it possible to monitor the shift from even-aged forest management to spontaneous forest development. In general, spontaneous development was marked mainly by aggregated tree distribution in the middle and lower layer, and a shift from normal to reversed J-shaped dbh distribution. The total volume of deadwood in one of the unmanaged plots amounted to 48.10 m3/ha (9.5% of living wood volume). The use of adequate structural and spatial analysis as indicators of natural forest development are discussed.Downloads
References
Aguirre O., Hui G., Gadow K.V., Jiménez J., 2003. An analysis of spatial forest structure using neighbourhood-based variables. For Ecol Manage. 183, 137-145.
Atici E., Colak A.H., Rotherham I.D., 2008. Coarse dead wood volume of managed oriental beech (Fagus orientalis Lipsky) Stands in Turkey. Investigación Agraria: Sistemas y Recursos Forestales 17(3), 216-227.
BARNA M., MARUSÁK R., 2003. Diameter structure of a beech parent stand at shelterwood cut phase. Folia oecologica 30(2), 91-98.
Bílek L., Remes˘ J., Zahradník D., 2009. Natural regeneration of senescent even-aged beech (Fagus sylvatica L.) stands under the conditions of Central Bohemia. J For Sci 55(4), 145-155.
Christensen M., Hahn K., Mountford E.P., Ódor P., Standovár T., Rozenbergar D., Diaci J., Wijdeven S., Meyer P., Winter S., Vrska, T., 2005. Dead wood in European beech (Fagus sylvatica) forest reserves. For Ecol Manage 210, 267-282.
Clark P.J., Evans F.C., 1954. Distance to nearest neighbour as a measure of spatial relationship in populations. Ecology 35, 445-453. http://dx.doi.org/10.2307/1931034
Emborg J., Christensen M., Heilmann-Clausen J., 2000. The structural dynamics of Suserup Skov, a nearnatural temperate deciduous forest in Denmark. For Ecol Manage 126, 173-189.
Foster D.R., Orwig D.A., Mclachlan J.S., 1996. Ecological and conservation insights from reconstructive studies of temperate old-growth forests. Trends in Ecology & Evolution 11, 419-424. http://dx.doi.org/10.1016/0169-5347(96)10047-1
Füldner K., 1995. Strukturbeschreibung von Buchen-Edellaubholz-Mischwäldern (Describing forest structures in mixed beech-ash-maple-sycamore stands). Ph D Dissertation, Göttingen University, Cuvillier Verlag, Göttingen.
Gadow K.V., 1993. Zur Bestandesbeschreibung in der Forsteinrichtung (New variables for describing stands of trees). Forst und Holz 48, 601-606.
Gadow K.V., 1999. Waldstruktur und Diversität (Forest structure and diversity). Allg Forst-Jagdztg 170, 117-122.
Gini C., 1921. Measurement of inequality on income. Econ J 31, 22-43. http://dx.doi.org/10.2307/2223319
Green P., Peterken G.F., 1997. Variation in the amount of dead wood in the woodlands of the Lower Wye Valley, UK in relation to the intensity of management. For Ecol Manage 98, 229-238.
Hanewinkel M., 2004. Spatial patterns in mixed coniferous even-aged, uneven-aged and conversion stands. Eur J For Res 123, 139-155.
Hopkins B., Skellam J.G., 1954. A new method for determining the type of distribution of plant individuals. Ann Bot 18, 213-227.
korpel'S ., 1995. Die Urwälder der Westkarpaten. Gustav Fischer Verlag, Stuttgart-Jena-New York. 310 pp.
Korpel's ., Pen Áz J., Saniga M., Tesar V., 1991. Pestovanie lesa. Príroda, Bratislava. 465 pp.
Leibundgut H., 1982. Europäische Urwälder der Bergstufe. Haupt, Bern, Stuttgart. 308 pp.
Meyer P., Tabaku V., Lüpke B. von, 2003. Die Struktur albanischer Rotbuchen-Urwälder-Ableitungen für eine naturnahe Buchenwirtschaft. Forstwissenschaftliches Centralblatt 122, 47-58. http://dx.doi.org/10.1046/j.1439-0337.2003.02041.x
Motz K., Sterba H., Pommerening A., 2010. Sampling measures of tree diversity. For Ecol Manage 260(11), 1985-1996.
Mountford M.D., 1961. On E. C. Pielou's index of non randomness. J Ecol 49, 271-275. http://dx.doi.org/10.2307/2257261
Nagel T.A., Svoboda M., Diaci, J., 2006. Regeneration patterns after intermediate wind disturbance in an oldgrowth Fagus-Abies forest in southeastern Slovenia. For Ecol Manage 226, 268-278.
Oheimb G. Von, Westphal C., Tempel H., Härdtle W., 2005. Structural pattern of a near-natural beech forest (Fagus sylvatica) (Serrahn, North-east Germany). For Ecol Manage 212, 253-263.
Ohser J., 1983. On estimators for the reduced second moment measure of point processes. Mathematische Operationsforschung und Statistik 14, 63-71.
Paluch J.G., 2007. The spatial pattern of a natural European beech (Fagus sylvatica L.)-silver fir (Abies alba Mill.) forest: a patch mosaic perspective. For Ecol Manage 253, 161-170.
Penttinen A., Stoyan D., Henttonen H., 1992. Marked point processes in forest statistics. For Sci 38(4), 806-824.
Petrás R., Pajtík J., 1991. Sústava cesko-slovenskych Objemovy ch tabuliek drevín. Lesnicky c asopis 1, 49-56. PIELOU E.C., 1959. The use of point-to-plant distances in the study of the pattern of plant populations. J Ecol 47, 607-613.
Pielou E.C., 1961. Segregation and symmetry in two species populations as studied by nearest neighbour relations. J Ecol 49, 255-269. http://dx.doi.org/10.2307/2257260
Pokorny J., 1963. Vy voj a druhová skladba c ernokostelecky ´ch lesu° do konce 18. století, C˘ ást II. Vy voj a obraz druhové skladby revíru Bohumile, Vode rady a Oplany. Sborník lesnické fakulty Vysoké s koly zeme de lské v Praze 6, 31-107.
Pommerening A., 2002. Approaches to quantifying forest structures. For 75(3), 305-324.
Rademacher C., Neuert C., Grundmann V., Wissel C., Grimm V., 2001. Was charakterisiert Buchenurwälder? Untersuchungen der Altersstruktur des Kronendachs und der raümlichen Verteilung der Baumriesen in einem Modellwald mit Hilfe des Simulationsmodells BEFORE. Forstwissenschaftliches Centralblatt 120, 288-302. http://dx.doi.org/10.1007/BF02796101
Rozas V., 2006. Structural heterogeneity and tree spatial patterns in old-growth deciduous lowland forest in Cantabria, northen Spain. For Ecol Manage 185, 57-72.
Schnitzler A., Borlea F., 1998. Lessons from natural forests as keys for sustainable management and improvement of naturalness in managed broadleaved forests. For Ecol Manage 109, 293-303.
Schütz J.P., 1999. Close-to-nature silviculture: is this concept compatible with species diversity? For 72(4), 359-366.
Schütz J.P., 2002. Silvicultural tools to develop irregular and diverse forest structures. For 75(4), 329-337.
Shannon C., Weaver W., 1949. The mathematical theory of communication. The University of Illinois Press, Urbana and Chicago.
Srámek O., 1983. SPR Vode radské buc iny I. a II. Památky a príroda.
Standovár T., Kenderes K., 2003. A review on natural stand dynamics in beechwoods of East Central Europe. Appl Ecol Environ Res 1(1-2), 19-46.
Sterba H., 2008. Diversity indices based on angle count sampling and their interrelationships when used in forest inventories. For 81(5), 587-597.
Sterba H., Zingg A., 2006. Abstandsabhängige und abstandsunabhängige Bestandesstrukur-beschreibung. Allg Forst-Jagdztg 177, 169-176.
Stoyan D., Penttinen A., 2000. Recent applications of point process methods in forestry statistic. Statist Sci 15(1), 61-78.
Tabaku V., 1999. Struktur von Buchenwäldern in Albanien im Vergleich mit deutschen Buchen-Naturwaldreservaten und Wirtschaftswäldern. Cuvilier Verlag, Göttingen. 206 pp. PMid:10546101
UNECE/FAO, 2000. Forest resources of Europe, CIS, North America, Australia, Japan and New Zealand (Industrialized temperate/ boreal countries). UN-ECE/FAO Contribution to the Global Forest resources Assessment 2000. Main report. Geneva Timber and Forestry Study papers, No. 17, United Nations, New York and Geneva.
Wolf A., 2005. Fifty year record of change in tree spatial patterns within a mixed deciduous forest. For Ecol Manage 215, 212-223.
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